Well, I've decided to let them just be where they are. I guess it'll be a few months before I find out what they are .... and the missing 4th one has now reappeared. I have no plans to add any mechanical filtration, just let the plants and rock do it.
Although their heads are just the size of a pin head, it's amazing that they can find each other in this 60L tank, and exhibit schooling behaviour.